When Evan Spiegel and Miranda Kerr met for a first date, at a kundalini class at Golden Bridge Yoga in Los Angeles, Spiegel arrived early and sat at the front of the studio. Kerr walked in late (45 minutes late, Spiegel says) and took a place at the back, where she giggled out of nervous habit. “When I was a child, my mom called me Giggling Gert,” she says.

The relationship between the co-founder of Snap Inc. and the supermodel has never been a typical billionaire-meets-model nerd-fantasy pairing. He is seven years her junior...

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